
William (Ned) Friedman
Arnold Professor of Organismic
and Evolutionary Biology
Director of the Arnold Arboretum of Harvard University
Phone: 617-384-7744
E-mail: ned@oeb.harvard.edu
Office: Arnold Arboretum
125 Arborway
Boston, MA 02130
Lab Website: http://www.oeb.harvard.edu/faculty/friedman
My research program focuses on the organismic interfaces between developmental, phylogenetic and evolutionary biology. Within the past fifteen years, remarkable advances in the study of the phylogenetic relationships of plants have provided the raw materials for critical studies of character evolution. Armed with hypotheses of relationships among organisms, I seek to explore how patterns of morphology, anatomy and cell biology have evolved through the modification of developmental processes. My work is primarily focused on the origin and subsequent diversification of flowering plants, Darwin's "abominable mystery."
Recent Publications
Madrid, E.N. and W.E. Friedman. 2010. Female gametophyte and early seed development in Peperomia (Piperaceae). American Journal of Botany 97: 1-14. | pdf
Madrid, E.N. and W.E. Friedman. 2009. The developmental basis of an evolutionary diversification of female gametophyte structure in Piper and Piperaceae. Annals of Botany 103: 869-884. | pdf | commentary
Friedman, W.E. and K.C. Ryerson. 2009. Reconstructing the ancestral female gametophyte of angiosperms: insights from Amborella and other ancient lineages of flowering plants. American Journal of Botany 96: 129-143. | pdf
Friedman, W.E. 2009. The meaning of Darwin's "abominable mystery." American Journal of Botany 96: 5-21. | pdf
Holloway, S.J. and W.E. Friedman. 2008. Embryological features of Tofieldia glutinosa and their bearing on the early diversification of monocotyledonous plants. Annals of Botany 102: 167-182. | pdf
Friedman, W.E., S.C.H. Barrett, P.K. Diggle, V.F. Irish and L. Hufford. 2008. Whither plant evo-devo? Investigating the evolution of plant form: conceptual integration from the molecular to the ecological. New Phytologist 178: 468-471. | pdf
Friedman, W.E. 2008. Hydatellaceae are water lilies with gymnospermous tendencies. Nature 453: 94-97. | pdf
Friedman, W.E., E.N. Madrid, and J.H. Williams. 2008. Origin of the fittest and survival of the fittest: relating female gametophyte development to endosperm genetics. International Journal of Plant Sciences 169: 79-92. | pdf
Winther J.L. and W.E. Friedman. 2008. Arbuscular mycorrhizal associations in Lycopodiaceae. New Phytologist 177: 790-801. | pdf | commentary
Friedman, W.E. 2006. Sex among the flowers. Natural History 115 (9): 48-53.
Friedman, W.E. 2006. Embryological evidence for developmental lability during early angiosperm evolution. Nature 441: 337-340. | pdf
Friedman, W.E., and J.H. Williams. 2004. Developmental evolution of the sexual process in ancient flowering plant lineages. Plant Cell 16: S119-S132. | pdf
Friedman W.E., R.C. Moore and M.D. Purugganan. 2004. The evolution of plant development. American Journal of Botany 91: 1726-1741. | pdf